Source: MONTANA STATE UNIVERSITY submitted to NRP
WHEAT TILLERING, BIOMASS, AND SEED YIELD ENHANCEMENT VIA MANIPULATION OF TEOSINTE BRANCHED 1 FUNCTION
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
ACTIVE
Funding Source
Reporting Frequency
Annual
Accession No.
1030066
Grant No.
2023-67014-39613
Cumulative Award Amt.
$644,293.00
Proposal No.
2022-10246
Multistate No.
(N/A)
Project Start Date
Sep 1, 2023
Project End Date
Aug 31, 2027
Grant Year
2023
Program Code
[A1141]- Plant Health and Production and Plant Products: Plant Breeding for Agricultural Production
Recipient Organization
MONTANA STATE UNIVERSITY
(N/A)
BOZEMAN,MT 59717
Performing Department
(N/A)
Non Technical Summary
This project uses Teosinte Branched 1 (TB1) mutations to increase plant biomass and seed yield in elite wheat varieties. TB1 is a gene that influences tillering in cereal plants.There are few known natural TB1 alleles for wheat and none that vary significantly in function. Furthermore, tight linkage between TB1 and Reduced Height 1, a major height gene, has inhibited selection of new alleles. Chemically induced TB1 mutants have allowed us to test the impact of reduced TB1 dosage in fixed Rht backgrounds. Our preliminary results to dateindicate that reducing TB1 function increases productive tillers and plant biomass by >10%. In this project will create near isogenic lines (NILs) varying for TB1 alleles in spring, winter, and durum wheat standard and semi-dwarf backgrounds. We will then test all NILs across multiple locations, environments, and planting densities to identify TB1 allelic combinations best suited for growth in target environments. Our approach will maximize productive tillers and grain yield potential. All genotypes developed in this project will be transferred to wheat breeding programs and lead to yield enhanced varieties. This will benefit growers, the state, and national economy. Tillering potential is a fluid trait that can be quite resilient, with the ability to adapt to a changing climate. This is an ever-important quality when working toward US and global food security.
Animal Health Component
40%
Research Effort Categories
Basic
60%
Applied
40%
Developmental
0%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
20115491081100%
Goals / Objectives
This project uses Teosinte Branched 1 (TB1) mutations to increase plant biomass and seed yield in elite wheat varieties. There are few known natural TB1 alleles for wheat and none that vary significantly in function. Furthermore, tight linkage between TB1 and Rht, a major height gene, has inhibited selection of new TB1 alleles. Our preliminary results indicate that reducing TB1 function increases productive tillers and plant biomass by >10%. Here, we will create BC5F4 NILs varying for TB1 alleles in spring, winter, and durum wheat standard and semi-dwarf backgrounds. We then will test all NILs across multiple locations, environments, and planting densities to identify TB1 allelic combinations best suited for growth in target environments. Our approach will maximize productive tillers and grain yield potential. All genotypes developed in this project will be transferred to wheat breeding programs and lead to yield enhanced varieties. This will benefit growers, the state, and national economy. Tillering potential is a fluid trait that can be quite resilient, with the ability to adapt to a changing climate. This is an ever-important quality when working toward US and global food security.
Project Methods
Objective 1: Creation and initial characterization of NILs varying in TB1 function. This will be carried out in durum, spring, and winter wheat backgrounds.To create genotypes varying in TB1 function that wheat breeders can use to improve tillering potential within their breeding programs to enhance plant biomass and/or seed yield. In this objective we integrate TB1 variant alleles into elite MT germplasm. This proposal builds upon our results in whichBC1F2 derived EMS TB1 homozygous durum lines were characterized. Our results demonstrate that reducing TB1 function increases plant growth. We will create NILs varying in TB1 function in spring, winter, and durum wheat.Objective 2: Comprehensive field evaluation of isolines (NILs) carrying single and double TB1 variants. We will carry out yield tests of TB1 durum, spring, and winter wheatNILs at multiple locations with multiple seeding densities. We will carry out detailed plant measurements for each growing condition. We haveobserved that different TB1 conditioned responses under different environments. While TB1 mutations increased tillering in each environment it was not necessarily significant in all environments nor is the magnitude of difference likely to remain constant. Therefore, it is important to carry out yield testing under varying environments. This will allow us to target TB1 genotypes to specific environments and conditions to produce the ideal tillering and yield phenotype. This objective will identify optimal TB1 genotypes for target locations and will aid breeders in variety development.It is well established that plants may adjust tiller number based on plant density. Therefore, it is important to grow TB1 NILs under varying conditions and environments. For example, the double variant NIL may perform quite differently under dryland production in Conrad, MT compared to irrigated production in Bozeman, MT. The advantages of this project may manifest in two ways. First, biomass and/or seed yield may be improved, leading to increased yield. Second, regardless of growing conditions, we believe that seeding rates may be reduced- resulting in cost savings to growers. We will test our genotypes at multiple locations to examine the sustainability of TB1 conditioned tiller increases under varying environments.

Progress 09/01/23 to 08/31/24

Outputs
Target Audience:Plant geneticists, plant breeders, plant biologists, seed companies, and farmers are reached via publications and presentations. The project was presented in field day presentations in the state of MT and at the Crop Science of America Annual Meeting. The results also appeared in poster and oral presentationsat other regional and national meetings. Seed company representations andplant breeders from other states were reachedvia direct one on one communication and via publications and presentations. Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided?Grad students and research associate have participated in Montana State Research Center field days to present the project to growers. They have also attended and presented the project at national meetings such as the Crop Science meeting. How have the results been disseminated to communities of interest?Results presented via presentation at field days throughout the state of MT to wheat growers. The results have also been presented to wheat breeding programs via presentation at regional and national wheat improvement meetings and at the national Crop Science conference. What do you plan to do during the next reporting period to accomplish the goals?We have identifiedTB1 mutations in both hexaploid and durum wheats andhave integrated them into durum, spring, and winter wheat breeding populations to create new experimental lines within Montana wheat breeding programs that have high tillering potential. We will be conducting field trials using different plant density and fertility levels at several locations in MT. We hope also to collaborate on forage trials in MT and in the southern US where wheat is more often grazed during the winter.

Impacts
What was accomplished under these goals? We have been testing the impact of increased primary tiller number upon yield under different rainfall levels and different planting densities. To do this, we crossed mutated alleles of the gene Teosinte Branched-1 (TB1) into Montana adapted durum wheat, spring wheat, and forage winter wheat cultivars. Durum lines have been tested over the past three seasons in four different locations, while spring and winter wheat lines have been tested for one summer in one location. Our results demonstratethat TB1 mutations increase productive tiller number and seed size across all environments and varieties tested. If conditions are good for crop growth and development after the jointing stage, TB1 mutant lines also show increased grain yields of up to 16 percent, with no significant decrease in protein content. When conditions are poor, TB1 mutant alleles do not have any negative impact on grain yield or quality. Because of the increase in tiller number, these lines are also able to be planted at lower seeding rates while maintaining high yield potential. We will be carrying out additional greenhouse and field trials in subsequent years for spring, winter, and durum wheat.

Publications

  • Type: Conference Papers and Presentations Status: Published Year Published: 2023 Citation: Hale, C.O., J.M. Martin, A.C. Hogg, and M.J. Giroux. 2022. Improving Durum Yield by Optimizing Tillers. Montana Grain Growers Convention. November 30-December 1, 2022. Great, Falls, MT.