Source: UNIVERSITY OF MISSOURI submitted to NRP
III: SMALL: DATAG: HOW GENOTYPE, ENVIRONMENT AND MANAGEMENT INTERACT TO INFLUENCE ANIMAL SIZE: AN EVALUATION OF JAMES' INTERPRETATION OF BERGMANN'S RULE IN BOS TAURUS CATTLE
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
ACTIVE
Funding Source
Reporting Frequency
Annual
Accession No.
1024843
Grant No.
2021-67021-33448
Cumulative Award Amt.
$499,999.00
Proposal No.
2020-08975
Multistate No.
(N/A)
Project Start Date
Nov 1, 2020
Project End Date
Oct 31, 2025
Grant Year
2021
Program Code
[A7302]- Cyber-Physical Systems
Recipient Organization
UNIVERSITY OF MISSOURI
(N/A)
COLUMBIA,MO 65211
Performing Department
Animal Science
Non Technical Summary
Computer vision and data-driven approaches are used to enhance existing DNA and growth beef industry datasets (>150,592 cattle with 50 million DNA variants) with dense data from three-dimensional (3D) cameras (~5,000 cows, ~7 million points per cow) to answer 170-year-old questions in ecology and evolution (Bergmann's Rule). Using yield, environmental stress, and pathogen resistance, the project creates DNA predictions to help farmers select cattle that thrive in the environment at their ranch, ultimately improving the environmental efficiency of beef production. Identifying adapted, energy efficient cattle allows family farms and ranches across the USA to be more profitable and improves animal welfare, an important societal issue. In Objective 1, 3D computer vision is used to measure the surface area and volume of cattle to test if surface-area-to-volume ratio (SA:V) influences genetics for growth under cold stress or growth under high feed resources. Larger animals have a lower SA:V, allowing them to retain heat more efficiently in winter. One hypothesis suggests larger animals live at higher latitudes (further north in this study) because they use the lower SA:V to better deal with cold stress. An alternative hypothesis suggests that animals are larger at higher latitudes because more food is available when the animals are growing. This research creates two DNA predictions, one tailored to cold stress and the second tailored to high feed resources. Genetic merits of animals within each of these production contexts will be directly compared to their SA:V ratios. In Objective 2, this project tests the hypothesis that random models versus machine learning methods better predict various cattle traits (feed intake, fat reserves, body weight, etc.) using processed 3D point clouds and environmental data. The project uses random models typically used in DNA predictions, but rather than DNA variants as the predictors, the analysis uses the large 3D camera data as predictors.Besides advances in computer vision algorithms (e.g. Structure from Motion) and new approaches to apply data-driven machine learning to agriculture problems, this project provides a definitive test of James's Rule (intraspecific Bergmann's Rule) that animal size increases with latitude due to an increased ability to maintain body temperature with a smaller SA:V ratio. This is contrasted with Geist's Rule that body size increases due to advantages of larger animals to utilize seasonal burst of food at higher latitudes. Explicitly comparing direct SA:V measurements with precise climate-by-growth and resource-by-growth predictions, this project will answer which force drives evolution in body size across latitudes. This will be the first definitive test of Bergmann's rule. By comparing random models versus deep learning to predict a variety of beef cow production traits using 3D camera data, this project indicates whether random models can replace or clarify artificial intelligence in many contexts. This project creates new data recording and selection tools based on Internet of Things and 3D computer vision algorithms to address the pressing need to measure and improve efficiency in pasture, cow-calf production (a $64 billion industry). This enables convenient and affordable measurement of beef cow efficiency (a vital aspect of beef production that is not currently predicted).Partnering with the University of Missouri School of Journalism Strategic Communication capstone course provides strategic plans to educate a variety of audiences on the true environmental impact of cattle and how technology adoption decreases this impact. The three-pronged approach 1) educates lay audiences 2) delivers online educational outreach to farmers/ranchers and 3) provides traditional extension programming to farmers/ranchers through regional and national in-person presentations. The first prong will reach a national audience through cable television. The second and third prong will reach approximately 7,000 people per year. With the National Center for Applied Reproduction and Genomics, a "Data Science Best Practices" module is created to give veterinary and graduate students a primer on data-driven agriculture. Partnerships with industry create applied genetic evaluations to predict genetic, environment, and management interactions, which allow precision selection not currently possible with one-size-fits-all national evaluations. The cow scanning system will be commercialized and added onto existing commercial platforms that collect feed and water intake. The 3D data is used to predict cow efficiency. Collectively, these impacts improve the financial security of rural farmers.
Animal Health Component
33%
Research Effort Categories
Basic
34%
Applied
33%
Developmental
33%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
3033310108025%
3023310101020%
3073310108010%
3033310208025%
3033310202010%
3033310107010%
Goals / Objectives
This project has two main goals.Use high-throughput phenotyping and "big data" to better predict production efficiency.Use high-throughput phenotyping and "big data" to test rules of evolution and ecology.For big data to be effective in livestock industries, we do not need more data, we need more useful data. Livestock industries need information-rich data to predict economically important traits that are not currently measured. Livestock production also needs cost effective, accurate ways to predict traits that are expensive to measure. However, electronic, high-dimensional records of beef cattle traits do not currently exist. We will address these needs by creating a cattle body scanner using from Red-Green-Blue, thermal and time-of-flight (ToF) cameras to create three-dimensional data with heat and color measures (4D-RGB); extract phenotypes from 4D-RGB data; and apply machine learning to predict traits.ObjectivesObjective 1)The first objective is to test the hypothesis that surface-area-to-volume ratio (SA:V) influences genotype-by-environment effects in Bos taurus beef cattle production using phenotypes based on 4D-RGB data generated with novel cattle scanners. An alternative hypothesis is that SA:V influences genotype-by-management effects in beef production.In 1847, Carl Bergmann proposed that body size increases at higher latitudes. It is believed that Bergmann's Rule was originally intended for comparisons across species. However, James's Rule has been suggested as an appropriate name for comparisons of body size and latitude within a species. There are two possible mechanisms for the effect of surface area (SA) and volume (V) on production in domesticated cattle. First is the Heat Conservation Hypothesis. Larger animals have lower SA:V compared to smaller animals. This allows larger animals to more efficiently conserve heat. Interestingly, it may actually be adaptation to heat and humidity, rather than to cold, that is responsible for the relationship between body size and latitude. Second is the Resource Availability Hypothesis (also referred to as Geist's Rule or eNPP Rule). This hypothesis states that primary plant productivity during the growing season (referred to as eNPP by Huston and Wolverton) influences the growth and reproduction of animals. This seasonal surge of resource availability favors larger animals.Beef cattle represent a unique opportunity to test these two hypotheses.Beef cattle are raised across the U.S. with essentially no management interventions to limit the effects of heat and cold stress, allowing the maximum influence of genotype × environment effects.Resources available to cattle, especially in the growing phase, are controlled by farmers and ranchers. These tend to fall into two groups: high-inputs (high resource availability) and low-inputs (low resource availability).There are large phenotypic and genomic databases available for beef cattle.Further, we are creating a system to measure the SA and V of cattle in an inexpensive, high-throughput manner. We will collect ~6,475 of SA and V phenotypes, which has not previously been done in any animal species.Restated, Objective 1 will test two hypotheses of forces that drive body size ecology and evolution in animals. We will test if heat retention as a function of larger body size altering SA:V affects adaptation to local climates. This is James's Rule (intraspecific Bergmann's Rule) that body size increases with latitude. We also test if resource availability (genotype × management interaction) is associated with body size. This is Geist's Rule, that pulses in primary plant productivity direct growth in animals.Objective 2)In Objective 2 we will test if multimodal data extracted from RGB, thermal and ToF cameras can be used to predict several production traits (feed intake, body condition score [level of fat reserves], mature weight, hair shedding, foot and skeletal scores, udder scores, etc.), comparing random regression models and a novel deep learning approach using XAI techniques.The second objective is to test the hypothesis that random models produce the same prediction accuracy as artificial intelligence (AI) methods when a large number of features are used to predict quantitative traits. Random regression models, such as Ridge Regression/GBLUP, BayesB, and BSLMM, are often used in genomic prediction to estimate the genetic merit of animals through a supervised process involving thousands to millions of DNA features. The effect of each feature is reported from these statistical models, and there are many software implementations that can handle very large data sets. However, these random regressions do not take advantage of the added discriminant power that neighboring/correlated data points, complementary features, and other underlying, nonlinear properties that a large number of features can provide with the use of many pattern recognition and machine learning approaches, e.g. deep learning (DL). Moreover, through the use of explainable AI techniques such as those derived from topological data analysis (TDA), we will test when and why random regression can/cannot compete with machine learning, and using what set of high dimensional image data. Indeed, eXplainable AI (XAI) is a promising research area, and this proposed objective has the potential to provide never-seen parallels, insights and tools to be used in random regression, in current PR/ML approaches, or both, in order to better understand predictions.As a related pursuit of better understanding AI and random regression methods, we will also test if the SA:V predicts an animal's appetite and feed intake. Perhaps the greatest ecological niche of cattle and other ruminants is their capacity to utilize cellulose and other plant materials indigestible to mammalian enzymes. The ruminant digestive tract evolved from non-ruminant species. Thus, similar to non-ruminant species, intake in cattle is regulated, in part, by chemostatic mechanisms that sense oxidation of metabolic fuels. However, different from non-ruminants, intake of cattle can also be regulated by ruminal volume. Ruminal volume is closely related to body size in ruminants, and when rates of ruminal fermentation of cellulosic materials are limited then intake of cattle is more closely described by ruminal fill than by caloric density of diets. Cattle that are more spherical in shape, have lower SA:V, as spheres have the lowest SA:V. We hypothesize that more spherical shaped cattle have more digestive system capacity and will eat more fermentable feed before their appetite is satiated.
Project Methods
Because this is an interdisciplinary project combining robotic vision, computer science, nutrition, metabolism, genetics and genomics, a multitude of methods will be used.First, Time of Flight, infrared, and Red-Green-Blue cameras will be used to scan cattle. Robotic vision algorithms (such as Structure from Motion, Bundle Adjustment, etc.) will be used to create a three-dimensional point cloud of the animal, from which surface area and volume will be measured.Surface area is a key parameter to measure an animal's basal metabolic rate.We will use multivariate genomic predictions models (such as genomic BLUP) to create genotype-by-environment, genotype-by-management, and genotype-by-environment-by-management genetic predictions of yearling growth. We will also create genomic predictions of surface-area-to-volume ratio. We will compare surface-area-to-volume to these context-specific genomic predictions of growth and body size.We will use various machine learning and statistical measures to predict cattle traits from both the raw and converted three-dimensional camera data. Predicted traits will include feed intake, body condition score, hair shedding score, and foot structure.

Progress 11/01/23 to 10/31/24

Outputs
Target Audience:During this reporting period the target audiences were broader than previous years, because with a patent filled, we no longer have to be mindful of public disclosures. The target audiences for this reporting period areindustry professionals,technology investors, scientists from several disciplines, farmers and ranchers. Changes/Problems:We have finally resolved many of the previous problems we were facing. When imaging cattle in November of 2023, we had a major failure of our chute system. We redesigned the chute so that it was much heavier duty and did a better job of containing the animals as they move through the chute. This new chute has worked very well. We have hired a senior research scientist in the Decker Computational Genomics group to lead genetic analyses. This should correct many of the staffing issues we have had in the first 3 years of the grant. What opportunities for training and professional development has the project provided?A masters student attended two industry conferences, namely the Beef Improvement Federation Symposium and the Angus Genetics Inc. Imagine Conference. This allowed the student to network with industry professionals and other scientists. This also helped the student learn about issues and opportunties in livestock production. How have the results been disseminated to communities of interest?We have filed a patent describing our technology and approach. We have published research showing the accuracy and advantages of our imaging approach. We have given presentations to producer and industry audiences about how this technology more accurate describes the metabolism of animals and how we can use it to be more efficient as an industry. What do you plan to do during the next reporting period to accomplish the goals?In the next reporting period, we will image the approximately 900 mature cows owned by the University of Missouri multiple times. We will also image thousands of cattle from private ranches in the southeast and northwest of the United States. We will produce genotype-by-temperature and genotype-by-management predictions of growth traits in large (90,000 animal) industry datasets. We will use these genetic interactionsto compare with surface area and volume measures to assess which ecological forces influence the observations motivating Bergmann's Rule. We will predict production traits using the data from the RGB and 3D imaging of cattle.

Impacts
What was accomplished under these goals? During this reporting period, we began to collect 3D images, surface areas, and volumes of large numbers of cattle. At University of Missouri Agriculture Experiment Station farms, we have collected data on several hundred cattle. We have also collected 3D images and indirect calorimetry measures for 20 steers at four time points along their growth curves. These data allow us to have more precise and generalizable estimates of the relationship between surface area, volume, and metabolic rate. We have begun to investigate genotype-by-environment and genotype-by-management effects in large beef cattle industry data sets. We have scored phenotypes of cattle that we can predict with machine learning models usingthe image data we have collected at the same time.

Publications

  • Type: Peer Reviewed Journal Articles Status: Published Year Published: 2024 Citation: Omotara, Gbenga, Seyed Mohamad Ali Tousi, Jared E. Decker, Derek Brake, and Guilherme N. DeSouza. "High-Throughput and Accurate 3D Scanning of Cattle Using Time-of-Flight Sensors and Deep Learning." Sensors 24, no. 16 (2024): 5275.
  • Type: Peer Reviewed Journal Articles Status: Published Year Published: 2024 Citation: Durbin, Harly J., Helen Yampara-Iquise, Troy N. Rowan, Robert D. Schnabel, James E. Koltes, Jeremy G. Powell, and Jared E. Decker. "Genomic loci involved in sensing environmental cues and metabolism affect seasonal coat shedding in Bos taurus and Bos indicus cattle." G3: Genes, Genomes, Genetics 14, no. 2 (2024): jkad279.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2024 Citation: Decker, Jared E. "Modern Research and Modern Tools to Match Cattle Genetics to the Environment" Beef Improvement Federation Symposium, Knoxville, Tennessee, June 11, 2024.


Progress 11/01/22 to 10/31/23

Outputs
Target Audience:During this reporting period the target audiences were industry professionals and technology investors. Changes/Problems:Many of the same issues that we struggled with in the previous reporting period continue to be an issue. We continue to struggle to recruit a postdoctoral fellow to join the group. This limits our ability to perform the research as quickly and efficiently as necessary. The animal science graduate student involved with the project took an industry position as the director of breed improvement for a beef breed association in March of 2023. We also continue to trouble shoot various technical and logistic issues with creating a 3-D cattle imaging system. What opportunities for training and professional development has the project provided? Nothing Reported How have the results been disseminated to communities of interest?We continue to have conversations with private firms about the opportunity to commercialize this technology. What do you plan to do during the next reporting period to accomplish the goals?We will have a first major data collection in November of 2023. We have started the process of hiring a senior research scientist to increase the speed of data analysis. We have recruited a masters student to help with collection of subjective cattle scores (body condition scores, foot scores, udder scores, hair scores, etc.). That student will begin in January of 2024.

Impacts
What was accomplished under these goals? We continued to refine and callibrate our 3D measurements. We trained a deep learning model that automatically separates pixels belonging to cattle from the background. This processes substantially increases the speed at which we can measure the surface area and volume of cattle, and most importantly, removes the need for manual intervention.

Publications

  • Type: Peer Reviewed Journal Articles Status: Published Year Published: 2023 Citation: Arisman, Brian C., Troy N. Rowan, Jordan M. Thomas, Harly J. Durbin, William R. Lamberson, David J. Patterson, and Jared E. Decker. "Evaluation of Zoetis GeneMax Advantage genomic predictions in commercial Bos taurus Angus cattle." Livestock Science 274 (2023): 105266.
  • Type: Peer Reviewed Journal Articles Status: Published Year Published: 2023 Citation: Grohmann, Caleb J., Caleb M. Shull, Tamar E. Crum, Clint Schwab, Timothy J. Safranski, and Jared E. Decker. "Analysis of polygenic selection in purebred and crossbred pig genomes using generation proxy selection mapping." Genetics Selection Evolution 55, no. 1 (2023): 62.


Progress 11/01/21 to 10/31/22

Outputs
Target Audience:During this reporting period the target audiences were industry professionals and technology investors. Changes/Problems:We continue to struggle to recruit a postdoctoral fellow to join the group. This limits our ability to perform the research as quickly and efficiently as necessary. The animal science graduate student involved with the project took an industry position as the director of breed improvement for a beef breed association. We also continue to trouble shoot various technical and logistic issues with creating a 3-D cattle imaging system. What opportunities for training and professional development has the project provided?An animal science graduate student supported by the project completed an internship at Angus Genetics Inc during the summer of 2021. This graduate student also attended the Gordon Research Seminar for early career trainees and Gordon Research Conference on Quantitative Genetics and Genomics. An engineering graduate student supported by the project completed an internship at EquipmentShare, where he collaborated on the development and integration of a vision system for safety in construction vehicles. This student also attended the14th International Conference on Advances in Quantitative Laryngology,Voice and Speech Research, where he won the Best Poster Award. How have the results been disseminated to communities of interest?Public disclosures regarding our 3-D cattle imaging were limited, as we were working with the University of MissouriTechnology Advancement Office to seek industry partners and a potential patent application. We have held several meetings with companies to discuss the commercial applications of our technology for livestock managment and genetic selection. The effects of genotype-by-environment interactions were discussed with farmers and ranchers at various beef industry meetings. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

Impacts
What was accomplished under these goals? We were able to find a suitable camera system for imaging cattle and reconstructing their 3-D shapes. We also identified the timing needed to properly synchronize these infrared cameras to avoid interference. We created an intial prototype of the 3-D imaging, including a chute. We imaged several dozen animals, including animals with both 3-D images, hide surface area measurements, and indirect calorimetry basal metabolic rate measurements. These data allow us to estimate the basal metabolic rate as a function of surface area and/or volume, which we have shown is more accurate than predictions based on the weight of the animal. Preliminary analyses on genotype-by-environment and genotype-by-management genetic interactions were completed.

Publications

  • Type: Journal Articles Status: Published Year Published: 2021 Citation: Keith, Monica H., Mark V. Flinn, Harly J. Durbin, Troy N. Rowan, Gregory E. Blomquist, Kristen H. Taylor, Jeremy F. Taylor, and Jared E. Decker. "Genetic ancestry, admixture, and population structure in rural Dominica." Plos one 16, no. 11 (2021): e0258735.
  • Type: Journal Articles Status: Published Year Published: 2022 Citation: Whitacre, Lynsey K., Mark L. Wildhaber, Gary S. Johnson, Harly J. Durbin, Troy N. Rowan, Peoria Tribe, Robert D. Schnabel et al. "Exploring genetic variation and population structure in a threatened species, Noturus placidus, with whole-genome sequence data." G3 12, no. 4 (2022): jkac046.
  • Type: Journal Articles Status: Published Year Published: 2022 Citation: Smith, Johanna L., Miranda L. Wilson, Sara M. Nilson, Troy N. Rowan, Robert D. Schnabel, Jared E. Decker, and Christopher M. Seabury. "Genome-wide association and genotype by environment interactions for growth traits in US Red Angus cattle." BMC Genomics 23, no. 1 (2022): 517. https://doi.org/10.1186/s12864-022-08667-6
  • Type: Conference Papers and Presentations Status: Published Year Published: 2022 Citation: Jared E. Decker. "Genomics 101 and More", Herefords in the Cove, Georgia Hereford Association, 2022.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2022 Citation: Jared E. Decker. "Population Genomics of Sustainable Livestock Production" 42nd ADSA Discovery Conference, 2022.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Jared E. Decker "Genotype-by-Environment-by-Management Interactions to Select More Sustainable Cattle." Bair Ranch Lecture, Department of Animal and Range Sciences, Montana State University. 2021
  • Type: Conference Papers and Presentations Status: Published Year Published: 2022 Citation: Jared E. Decker. "Combining Quantitative Genetics and Population Genomics to Improve Beef Sustainability" Purdue University 2022.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2022 Citation: Jared E. Decker "Combining population genomics and quantitative genomics to improve livestock sustainability" AGBT-Ag, The Genome Partnership, 2022.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2022 Citation: Knowles A.J., Lu, Duc, Garcia, Andre, Miller, Steven, Retallick, Kelli, Decker, Jared E. Comparisons of Genetic Predictions for Heifer Pregnancy, First Service Pregnancy, and Service Number in American Angus Heifers. Gordon Research Conference on Quantitative Genetics and Genomics. Februrary 2022.


Progress 11/01/20 to 10/31/21

Outputs
Target Audience:The improvements to Surface from Motion and Bundle Adjustment will be targeted to robotic vision and computer science audiences. Our surface area, volume, and trait predictions using machine learning methods will be targeted towards scientists with interests in high-throughput phenotyping and big data. Changes/Problems:Typical issues with prototype development were encountered. However, we were surprised to find the Intel's RealSense sensor created a distorted and wavey point cloud. This necessitated switching to a different brand of sensor. What opportunities for training and professional development has the project provided?Three graduate students have been mentored as part of the project. They have been deeply involved in the planning and implementation of the research. How have the results been disseminated to communities of interest?Public disclosure of the technology has been intentionally limited to maintain our ability to potentially patent our device and approach. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

Impacts
What was accomplished under these goals? In 2020 to 2021, we created our first 3-D cattle scanner prototype. We scanned two steers. After slaughtering the animals, we could compare the animal's surface area from the computer vision model to the surface area from the animal's hide. Initial computer scripts to capture, processes, and store the images were written.

Publications

  • Type: Journal Articles Status: Published Year Published: 2021 Citation: Mabry, Makenzie E., Troy N. Rowan, J. Chris Pires, and Jared E. Decker. "Feralization: confronting the complexity of domestication and evolution." Trends in Genetics 37, no. 4 (2021): 302-305.
  • Type: Journal Articles Status: Published Year Published: 2021 Citation: Braz, Camila U., Troy N. Rowan, Robert D. Schnabel, and Jared E. Decker. "Genome-wide association analyses identify genotype-by-environment interactions of growth traits in Simmental cattle." Scientific reports 11, no. 1 (2021): 13335.
  • Type: Journal Articles Status: Published Year Published: 2021 Citation: Rowan, Troy N., Harly J. Durbin, Christopher M. Seabury, Robert D. Schnabel, and Jared E. Decker. "Powerful detection of polygenic selection and evidence of environmental adaptation in US beef cattle." PLoS Genetics 17, no. 7 (2021): e1009652.
  • Type: Journal Articles Status: Published Year Published: 2021 Citation: Crum, Tamar E., Robert D. Schnabel, Jared E. Decker, and Jeremy F. Taylor. "Taurine and indicine haplotype representation in advanced generation individuals from three American breeds." Frontiers in Genetics 12 (2021): 758394.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2020 Citation: Decker JE (2020) Matching Genetics to Environment Using Genomics Synthesis of results from USDA-NIFA Food Security Grant on Local Adaptation in Beef Cattle. The 6th International Conference of Quantitative Genetics , November 10, 2020. (Oral Presentation)
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Decker JE (2021) Promising Projects in the Pipeline. Ag Innovation Forum, The Agricultural Business Council of Kansas City, February 18, 2021 (Panel Discussion)
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Decker JE, (2021) Combining Quantitative Genetics and Population Genomics to Improve Beef Sustainability. ISAG 2021, International Society for Animal Genetics, July 26, 2021 (Plenary Presentation)
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Decker JE, de Leon N (2021) Grand Challenge: Digital Agriculture/Smart Farms. North Central Mini Land-Grant Meeting, August 2, 2021 (Oral Presentation)
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Decker JE. (2021) Combining Quantitative Genetics and Population Genomics to Improve Beef Sustainability. University of Guelph Centre for Genetic Improvement of Livestock.
  • Type: Conference Papers and Presentations Status: Published Year Published: 2021 Citation: Decker JE. (2021) Genotype-by-Environment-by-Management Interactions to Select More Sustainable Cattle. Bair Ranch Lecture, Department of Animal and Range Sciences, Montana State University.