Source: CORNELL UNIVERSITY submitted to NRP
FLY MANAGEMENT IN ANIMAL AGRICULTURE SYSTEMS AND IMPACTS ON ANIMAL HEALTH AND FOOD SAFETY
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
1023341
Grant No.
(N/A)
Cumulative Award Amt.
(N/A)
Proposal No.
(N/A)
Multistate No.
S-1076
Project Start Date
Oct 1, 2020
Project End Date
Sep 30, 2023
Grant Year
(N/A)
Program Code
[(N/A)]- (N/A)
Recipient Organization
CORNELL UNIVERSITY
(N/A)
ITHACA,NY 14853
Performing Department
Entomology
Non Technical Summary
House flies are major pests of animal production facilities as they transmit numerous diseases and greatly reduce productivity. Insecticides are commonly used for controlling this pest, but resistance is a major problem. This project will monitor insecticide resistance levels, identify the genes underlying resistance, and evaluate new insecticides in order to inform future control strategies.
Animal Health Component
50%
Research Effort Categories
Basic
50%
Applied
50%
Developmental
(N/A)
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
72131101130100%
Knowledge Area
721 - Insects and Other Pests Affecting Humans;

Subject Of Investigation
3110 - Insects;

Field Of Science
1130 - Entomology and acarology;
Goals / Objectives
New technologies for management of biting and nuisance flies in organic and conventional systems a. Novel push-pull strategies (NE, NC, USDA-NE, USDA-FL)
b. Evaluation of improved monitoring systems (USDA-NE, CA, TN, NM)
c. Novel toxicants, biopesticides, and delivery systems (TX, USDA-FL, USDA-NE, FL, NE, PA, NM)
d. Non-pesticide management options (mechanical) (FL, NC, NE, USDA-NE, USDA-FL, USDA-TX, PA, TN)
Insecticide resistance detection and management a. Assessment of insecticide resistance (TX, NY, USDA)
b. Leveraging the Stomoxys and Musca genomes for novel control measures (NY, USDA)
Project Methods
1. We have extensive experience evaluating resistance in house fly populations and will use the methods we have established (diagnostic concentrations) to evaluate the levels of resistance to insecticides. A member of our S1076 group has put together an online list of all the insecticides that are labeled for fly control in each state. Collaborators will identify "hot spots" (i.e. facilities having fly control problems), collect flies, send pupae and we (CU, PSU, NIU) will test resistance levels to the insecticides available in that state. We anticipate doing this at three or four geographically widespread sites.2. No new insecticides have been registered as premise sprays for house fly control in over 20 years, despite numerous new insecticides being discovered. This leaves a number of recently developed insecticides with potential for fly control. Working with collaborators in our S1076 group, including government and industry representatives, we will select ten insecticides that are collectively viewed as having the greatest potential for house fly control. We (NIU, Cornell, USDA) will examine the toxicity of these compounds to susceptible and resistant strains, to evaluate efficacy and the potential for cross-resistance, respectively.3. Using the recently collected KS17 strain of house flies that is no longer controlled with pyrethroid insecticides we will seek to identify the mutations responsible for resistance using a technique called bulk segregant analysis (BSA) as follows. Unmated KS17 strain females will be selected at a dose of 800 μg/fly to insure it is homozygous for the resistance alleles. KS17 will be crossed to a susceptible strain and the F1 flies will be allowed to interbreed. This process will continue until the F7. Flies from the F7 will be split into two groups. One will be unselected and one will be selected at 800 μg/fly (a dose that allows only homozygous resistant individuals to survive). We will sequence the genomes of the two parental strains, the unselected F7 and the F7 survivors. A BSA will be carried to identify the loci responsible for resistance. The resolution of the resistance locus will be improved using amplicon sequencing. These analysis is facilitated by the completed house fly genome and recent long read sequencing. The genes present at each locus will be evaluated for their potential role in resistance and validation experiments will be carried out. The PI has all the necessary skill sets to complete these analyses. 4. In the last year of our study, flies will be collected from multiple locations in NY and throughout the USA (by the PI and by collaborators in the multistate project). We anticipate making about 14 collections from across the USA. We (Cornell, NIU and PSU) will determine the levels of resistance to the seven most commonly used insecticides and we (Cornell) will determine the frequencies of the resistance mutations.

Progress 10/01/20 to 09/30/21

Outputs
Target Audience: Nothing Reported Changes/Problems:The pandemic caused numerous problems. We had delays in fly collections, we had a shortage of labor (COVID restrictions) and a major facilities problem (our house fly rearing facility was broken for 3 months). All of this made for a very challenging year in which it was hard to make research progress. What opportunities for training and professional development has the project provided? Nothing Reported How have the results been disseminated to communities of interest? Nothing Reported What do you plan to do during the next reporting period to accomplish the goals?We will conduct one more round of fluralaner selection to generate what we anticipate will be a homozygous resistant strain. We will examine the effects of different synergists on fluralaner resistance to gain insights into the mechanisms of resistance. We will determine the inheritance of the resistance. We will extract DNA for individual flies from all of the collections and genotype them to determine the frequencies of the susceptible, kdr, super-kdr, kdr-his, 1B and Type N alleles. We will then prepare the results for publication.

Impacts
What was accomplished under these goals? A nationwide collection of house flies was undertaken in 2021. House flies were obtained from 19 different locations. These populations were used for two purposes: to select a strain of flies resistant to fluralaner and to determine the frequencies of the different Voltage gated sodium channel (Vgsc) mutations that were present. After three selections we have already obtained a highly resistant strain. This strain will be used to characterize the resistance found. We have begun our analyses of the Vgsc alleles and anticipate completing these analyses in 2022.

Publications