Source: UNIV OF PENNSYLVANIA submitted to NRP
THE EFFECTS OF COLONIZATION WITH CLOSTRIDIOIDES DIFFICILE ON THE FECAL MICROBIOME OF DAIRY CALVES
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
1021792
Grant No.
(N/A)
Cumulative Award Amt.
(N/A)
Proposal No.
(N/A)
Multistate No.
(N/A)
Project Start Date
Jan 1, 2020
Project End Date
Sep 30, 2021
Grant Year
(N/A)
Program Code
[(N/A)]- (N/A)
Recipient Organization
UNIV OF PENNSYLVANIA
(N/A)
PHILADELPHIA,PA 19104
Performing Department
CLINICAL STUDIES
Non Technical Summary
Calf diarrhea is a major cause of illness and death in dairy calves, and it represents a major economic loss for the dairy farmer. Many factors can contribute to the development of calf diarrhea, including management practices on the farm and animal nutrition, but ultimately this disease is caused by the flourishing of pathogenic bacteria in the animal's gut that overwhelm the "good bacteria" that keep the intestinal tract healthy. Studies have shown that the bacterial make-up of the calf gut (the "microbiome") is critical to calf health.One particular microbe that can be found in young calves is called Clostridioides difficile, or "C. diff". In people and in certain species of animals, C. diff can cause debilitating and sometimes life-threatening diarrhea. One of the ways it does this is by outcompeting the native flora of the gut and producing toxins that damage the lining of the intestine. It is unclear whether C. diff can cause diarrhea in calves. However, the presence of C diff alone, even at levels insufficient to cause diarrhea, has been shown to be associated with an altered gut microbiome in people and in several species of animals. We recently demonstrated such a finding in puppies, and we now intend to explore whether similar associations between C. diff and the gut microbiome occur in dairy calves. We propose to compare the gut microbiome of dairy calves that carry C. diff and calves that do not. Having a better understanding of the factors that contribute to a healthy gut can eventually lead to better control of calf diarrhea and the prevention of illness and death in calves and economic loss for the farmer.
Animal Health Component
0%
Research Effort Categories
Basic
100%
Applied
0%
Developmental
0%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
71234101110100%
Goals / Objectives
Infectious diarrheal disease is one of the main causes of mortality in dairy calves (1, 2), and calves less than 30 days of age are at highest risk of developing diarrhea (3, 4). Studies have shown that gut microbial composition is associated with gut health and the likelihood of diarrhea: reductions in microbial diversity are associated with an increased incidence of diarrhea (5), and the colonization of the calf gut with beneficial bacteria along with the decreased colonization of potential pathogens decreases the likelihood of calf diarrhea (6). Clostridioides difficile is a spore-forming anaerobic, gram-positive bacillus that is a significant enteric pathogen in many species of animals. Colonization with C. difficile has been shown to be associated with reduced gut microbial diversity and increased colonization of pathogenic bacteria in people (7, 8), and we recently demonstrated a similar association in puppies (9). Dairy calves, like the neonates of other species, are colonized with C. difficile at high rates, with reported prevalences ranging from 28-56% (10, 11). While there is some evidence that infection with C. difficile can result in diarrhea in calves (12), the effect of the asymptomatic colonization of calves on the gut microbiome is unknown. Given the crucial role of the gut microbiome in providing colonization resistance against pathogens that cause diarrhea (13, 14), a better understanding of the effect of pathogens such as C. difficile on the calf gut microbiome is needed. The goal of this study is to define the gut microbiota features associated with C. difficile colonization in dairy calves and to define the effects of calf age, diet, and farm on the risk of colonization.
Project Methods
Methods:We are currently in possession of a collection of 109 fecal samples from calves less than 14 days of age from 20 dairy farms in Pennsylvania. Pooled samples from each farm have undergone anaerobic culture for C. difficile as previously described (9). We propose to individually culture samples from the pooled samples that previously tested positive (n=36) and to submit all fecal samples (n=109) for 16S sequencing. Briefly, DNA will be extracted from the fecal samples, and the V4 region of the 16S rRNA gene will be amplified using barcoded primers for use on the Illumina platform (15). Sequencing will be performed using 250-base paired-end chemistry on an Illumina MiSeq instrument at the Center for Host Microbial Interactions at the University of Pennsylvania School of Veterinary Medicine.The effects of calf age, farm, and diet on culture status will be analyzed by logistic regression. Metrics of alpha and beta diversity of the fecal microbiota will be calculated using the qiime diversity core-metrics-phylogenetic function in qiime2 and visualized using QIIME2 and Emperor (16). The relative contributions of different microbial taxa that characterize the differences between C. difficile culture positive and negative calves will be assessed through linear discriminant analysis effect size (LEfSe).

Progress 01/01/20 to 09/30/21

Outputs
Target Audience:Stakeholders of calf health, including producers and veterinarians, represent the target audience for this study. The findings will be published in Plos One, a journal with a large veterinary audience.? Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided?This project provides insight into the microbiome of dairy calves during a critical period of development. The knowledge acquired in this study will be useful for understanding factors that modulate the gut microbiome and potential risk factors for neonatal diarrhea. How have the results been disseminated to communities of interest?The results of the study will be published in an upcoming edition of the Plos One journal, which has a large audience of scientists and veterinarians. The principal investigator will also present the findings of the study at regional and national scientific meetings. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

Impacts
What was accomplished under these goals? This project is now complete, and the associated manuscript has been accepted for publicationinPlos One. In this study, we sought to define microbial features associated with C. difficile colonization in pre-weaned dairy calves less than 2 weeks of age. We characterized the fecal microbiota of 80 calves from 23 different farms using 16S rRNA sequencing and compared the microbiota of C. difficile-positive (n=24) and C. difficile-negative calves (n=56). Farm appeared to be the greatest source of variability in the gut microbiota. When controlling for calf age, diet, and farm location, there was no significant difference in Shannon alpha diversity (P= 0.50) or in weighted UniFrac beta diversity (P=0.19) between C. difficile-positive and -negative calves. However, there was a significant difference in beta diversity as assessed using Bray-Curtiss diversity (P=0.0077), and C. difficile-positive calves had significantly increased levels of Ruminococcus (gnavus group) (Adj. P=0.052), Lachnoclostridium (Adj. P=0.060), Butyricicoccus (Adj. P=0.060), and Clostridium sensu stricto 2 compared to C. difficile-negative calves. Additionally, C. difficile-positive calves had fewer microbial co-occurrences than C. difficile-negative calves, indicating reduced bacterial synergies. Thus, while C. difficile colonization alone is not associated with dysbiosis and is therefore unlikely to result in an increased likelihood of diarrhea in dairy calves, it may be associated with a more disrupted microbiota.

Publications


    Progress 01/01/20 to 09/30/20

    Outputs
    Target Audience:Study not finished - no stakeholders reached yet Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided?A veterinary student was involved in this project and assisted with data collection and analysis. How have the results been disseminated to communities of interest? Nothing Reported What do you plan to do during the next reporting period to accomplish the goals?Complete data collection and analysis as well as manuscript redaction.

    Impacts
    What was accomplished under these goals? The study was not yet completed at the time of submission of this report.

    Publications