Source: NORTH CAROLINA A&T STATE UNIV submitted to NRP
DIVERSIFYING TREE HOST RANGE FOR SUSTAINABLE PRODUCTION OF BIANCHETTO TRUFFLES- TUBER BORCHII
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
1019921
Grant No.
(N/A)
Cumulative Award Amt.
(N/A)
Proposal No.
(N/A)
Multistate No.
(N/A)
Project Start Date
Oct 1, 2019
Project End Date
Sep 30, 2022
Grant Year
(N/A)
Program Code
[(N/A)]- (N/A)
Recipient Organization
NORTH CAROLINA A&T STATE UNIV
1601 EAST MARKET STREET
GREENSBORO,NC 27411
Performing Department
Natural Resources & Environmental Design
Non Technical Summary
Truffle farming is an emerging agricultural system in the US. Efforts to develop sustainable systems for its cultivation are critical and needed from the onset. Truffles develop in association with roots of live trees in a symbiotic relationship. As of today, truffles are the most expensive agricultural produce depending on the species ($200-$1000/lb) and will be a big boost to US farmers if developed sustainably. Approximately 95% of truffle consumed in the US are imported from Europe. Their increased global demand and decline in collection from the wild, has resulted in proliferation of cultivated truffle orchards. The Mushroom Biology & Fungal Biotechnology Laboratory at North Carolina A&T State University Greensboro is at the forefront of domesticating both local and European truffles in the US. Significant progress has been made in the domestication of Tuber borchii (common name = bianchetto truffle) on just one host plant- loblolly pine. Expanding the host range of Tuber borchii to other economically important and ecologically diverse pine species will potentially increase its production and sustainability in the South East USA. The first objective is to study and diversify the host range and ecological niche for T. borchii truffles. Accumulated from several years of studies on truffle synthesis on loblolly pine seedlings, we have observed substantial reduction in seedling growth due to nutrient limitations. The dynamic balance of nutrient supply to support optimal seedling growth, mycelia growth, mycorrhizal synthesis, and root colonization is not known for T. borchii. The second objective will focus on determining the optimum level of inorganic nutrients (nitrogen, phosphorus and potassium (NPK)), which will most enhance pine seedling growth and colonization by T. borchii in the greenhouse.To achieve these objectives, three pine species- Long leaf pine (Pinus palustris), short leaf pine (Pinus echinata) both agroforestry trees, and dwarf pine (Pinus strobus) an ornamental tree will be mycorrhized with Tuber borchii and their performance will be compared to the already-established truffle host tree- the loblolly pine. In the second part of the project, seedlings of loblolly pine will be mycorrhized and exposed to different levels of NPK fertilizer. Plant growth parameters and truffle colonization will be monitored and compared to non-fertilized controls. Mycorrhized trees from selected fertilizer treatments will be out-planted to the field and monitored throughout the project.It is expected that the three pine species will be suitable hosts for truffle colonization, establishment and eventual fructification at maturity. However, the extent to which each species support mycorrhization will be determined by experimentation. NPK fertilizer will definitely enhance plant growth but we will be able to determine levels that will concurrently enhance plant growth and truffle symbiosis. The project will have huge impact on increasing truffle production in the US by 1) diversifying the host range of T. borchii truffles from one to four trees, 2) increasing the regions where truffles could be cultivated since these species grow in different regions and ecological niches in the US, and 3) publishing data that will be useful for truffle growers and stakeholders.
Animal Health Component
30%
Research Effort Categories
Basic
20%
Applied
30%
Developmental
50%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
2051470110250%
1310699106025%
1251470110025%
Goals / Objectives
The overall goal of the project is to make truffle production more sustainable and wide spread in the entire US. The goal will be met with the following objectives:To evaluate three Pinus species as host for mycorhhizae and cultivation of Bianchetto truffles (Tuber borchii)To evaluate the effect of fertilizer application on mycorrhizae formation in greenhouse and their survival in field plots
Project Methods
Objective 1. To evaluates three Pinus species as host for mycorhhizae and cultivation of Bianchetto truffles (Tuber borchii)Three pine species (Long leaf pine (Pinus palustris), Short leaf pine (Pinus echinata), dwarf pine (Pinus strobus)) will be evaluated for their ability to support mycorrhizae formation with the T. borchii. Seeds from these pine species will be stratified, germinated and infected with T. borchii. Data will be collected on plant growth and development as well as persistence and colonization of pine roots by truffle fungus through physical, microscopic, and molecular techniques. Mycorrhization results obtained will be compared among the test species and with loblolly pine (Pinus taeda) which we have demonstrated to be a suitable host for T. borchii. Objective 2. To evaluate the effect of fertilizer application on mycorrhizae formation and their survival in field plots.There are conflicting results on the effect of fertilizer on mycorrhizae formation and progression. This objective will provide a first-hand information on how nutrient which are supposed to enhance seedling growth and development will influence mycorrhizae formation. Seeds of loblolly pine will be stratified, germinated, and infected with Tuber borchii fungus. Mycorrhized trees will be treated with and without NPK fertilizer at 4 levels (0, Low, Medium, and High) in the greenhouse. Host plant growth and development, nutrient uptake, and efficiency of mycorrhization will be measured. Seedlings from selected fertilizer treatments will be outplanted in the field and monitored. Seedling growth and development, truffle survival in pine root, and changes in microbial communities in orchards will be monitored.

Progress 10/01/19 to 09/30/20

Outputs
Target Audience:Diversifying pine host is one of the strategies to make truffle production sustainable. In North Carolina which has the most planted acreage of truffle orchards, only one pine species has effectively been used as host to synthesize truffles. Increasing the truffle pine host species would increase the ecological niches of truffles both locally and nationwide. The project targets truffle farmers- who will have a diversity of tree host to grow their truffle, truffle seedling producers- who are seriously looking for ways to enhance truffle seedling production and reduce time to seedling out-planting to field, and the truffle industry- which is still at a nascent stage. Also of interest is the opportunity that the data will give to extension agents to develop training manuals and deliver extension services to truffle farmers. Changes/Problems:The project will be needing extension of time due to delays caused by COVID-19. What opportunities for training and professional development has the project provided?Part of our approach to knowing whether truffles thrive in the presence of fertilizer is to quantify their abundance in soil. Project team has the opportunity to acquire more skills in the area of genomics. The PI attended a genomics training in Sept, 2020. Opportunities for training have largely been stalled by COVID-19. How have the results been disseminated to communities of interest? Nothing Reported What do you plan to do during the next reporting period to accomplish the goals?Several activities are planned for the next reporting cycle. The plant growth parameters and nutrient uptake will be fully analyzed. We will develop and validate the RT-PCR protocol for truffle DNA quantification. We will also complete the germination and study of the growth patterns of Pinus strobus, Pinus echinata and Pinus palustris, which is objective 1.

Impacts
What was accomplished under these goals? Research conducted so far focused on evaluating the effect of fertilizer application on pine truffle mycorrhizae formation and pine seedling health in the greenhouse. The goal is to know if fertilizer application which is known to enhance pine growth will have any effect (good or bad) on mycorrhizae synthesis, persistence and establishment in orchard soils. Results show that application of some level of fertilizer enhance pine growth as reported in literature. Data analyzed so far indicate that pine plants which are fertilized and mycorrhized with truffles, have more robust growth than the fertilized and non-mycorrhized plants as deduced from plant heights, root and shoot weights, as well as root:shoot ratio. Change in knowledge: Fertilizers are known to improve pine growth when applied at the right proportion. What is not known is whether fertilizer affect truffle mycorrhization of pine trees or truffle abundance in soil. Rudimentary data so far show that certain levels of fertilization could enhance pine growth in the presence of truffle mycorrhization. This project was largely affected by COVID, and expected output is not timely. We envisage that at the completion of this object, a baseline fertilizer regime for truffle cultivation will be established.

Publications