Source: UNIVERSITY OF TENNESSEE submitted to NRP
PRECISION FORESTRY: SILVICULTURAL METHODS FOR ARTIFICIAL REGENERATION AND THE ENHANCEMENT OF INTERMEDIATE STANDS IN HARDWOOD MANAGEMENT
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
1005809
Grant No.
(N/A)
Cumulative Award Amt.
(N/A)
Proposal No.
(N/A)
Multistate No.
(N/A)
Project Start Date
Mar 16, 2015
Project End Date
Jan 31, 2019
Grant Year
(N/A)
Program Code
[(N/A)]- (N/A)
Recipient Organization
UNIVERSITY OF TENNESSEE
2621 MORGAN CIR
KNOXVILLE,TN 37996-4540
Performing Department
Forestry, Wildlife & Fisheries
Non Technical Summary
Precision Forestry is a silvicultural system to regenerate hardwood forests by artificially regenerating small harvested openingswithsuperior seedlings and enhancing these trees at mid-rotation with crop tree treatments, including release and/or fertilization.Group tree selection openings can be created after any harvestand provide a value-addedalternative totraditional high grading with the ability to create regeration opportunities on areas with poor or understocked residual conditions. Small clearcuts traditionally regenerate tofast growing butless desirable, pioneer species such as sweetgum. sycamore and yellow poplar. By enriching the openings with highly desirable species, such as oak, and using seedlings withtheability to compete,theforest's ability to sustain a wider range of values, including timber, wildlife and recreational benefits,is greatly enhanced. Stand enrichment is the goal and seedlingsare planted on a wide spacing, generally 100/acre,with the expectation that not all will live to a free-to-grow condition.The guiding philosophy is 36 successful trees on-site at maturity is optimum, representing a signficant positiveinfluence and for some species, such as oak having large crowns, 36 trees canessentially capture the stand. At mid-rotation, a subset of the planted trees, i.e., those having demonstrated best potential for the trait deisred, can be selected for crop tree treatements.Studies are underway testing seedling survival and repsonse to crop tree treatments. In this study all seedlings are selected from among those destined as orchard trees in the develpment of hardwood seed orchards.This unique arrangement allows comparison of orchard and wild settings bewteen trees with known lineages.The Summary section has not changed. The only section that has changed is the budget section under the proposal tab. Please see page 14 of the updated proposal. REEport will not allow a project change when only the proposal has been changed.
Animal Health Component
90%
Research Effort Categories
Basic
10%
Applied
90%
Developmental
(N/A)
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
1230621107025%
1230699106075%
Goals / Objectives
Goals The objectives are focused on the successful establishment of artificial regeneration and improvement of the intermediate stages of southern hardwood stands. Quantitative demonstration of these parameters support the overarching goal: the development of a silvicultural stratagem specific to southern hardwoods and something that has a genuine chance of acceptance among private landowners. Pine silviculture can be distilled into a clearly recognized and applied set treatments. They are accepted. There is no birth-to-harvest set of treatments for hardwoods, especially where artificial regeneration is utilized outside of plantation management. With the availability of robust seedlings the research will develop a silvicultural package to address the unique needs of hardwoods where repeated harvests are applied to mature stands. Objectives1) develop a practicable methodology that can stand alone or can influence traditional landowner harvest tendencies, i.e., high-grading, with a regeneration strategy to improve species composition and forest health,2) evaluate the survival and productivity of enrichment plantings of oak (and other) species in group selection openings,3) examine the effects and interactions of site characteristics, including light, soil type, moisture and competition on highly selected, pedigreed seedlings within these gaps over time,4) examine the effects and interactions of release and fertilization treatments on the intermediate stand dynamics of both the planted trees and in natural stands,5) determine economic feasibility of these treatments, potential alternative strategies and ancillary effects (e.g., where crop trees are released, what effect is accomplished on trees in the "competition cluster" surrounding the crop tree). In all cases, these objectives cannot be achieved unless the studies are relatively long-term.
Project Methods
Precision Forestry: Artificial Regenration of Group Selection Openings Over 8,000 seedlings have been planted in 1-to-3 acre openings created in mature hardwood forests on the Ames Plantation in western Tennessee. Precision Description: Crop Tree Enhancement -- As an Example: Description of the Initial White Oak StudyInitially, and as a descriptive model for the whole study, a uniform, 40-year-old, single cohort, 20-acre hardwood stand on the Ames Plantation in western Tennessee was assessed for potential crop trees. The stand was divided into 720 cells measuring 35X35 feet. Within each cell a crop tree was selected, if present, on the basis of species, form class, and Crown Class Point System rating. Crop trees were available in 653 cells (91%). The stand was divided with a randomized complete block design with one acre equaling a treatment block of 36 trees.The stand was dominated by white oak and that species was primarily selected as crop trees. Species adequately represented to allow statistical testing were:Species Number of Crop TreesWhite Oak...................................445Southern Red Oak..........................101Black Oak................................... 38Scarlet Oak.................................. 33Black Cherry................................ 27TreatmentsThe treatments were Control (no treatment); Release (crown release on at least 3 sides); Fertilization (150 pounds per acre of available N applied as urea and 35 pounds per acre of Phosphorus applied as triple-super-phosphate) and Release and Fertilization.The Number of crop trees by species and treatment were as follows: Treatment ---------------------------------------------------------------------------------- Species Control Release Fertilization CombinationWhite Oak 104 117 122 102Southern Red Oak 26 28 21 26Black Oak 9 8 3 16Scarlet Oak 7 5 15 6Black Cherry 7 5 6 9These data were taken for all crop trees:Crop Tree Assigned a Number and Tagged at 5 feet above ground levelSpeciesDBH (taken 6inches below tag)Total HeightHeight to Green CrownBasal Area around the Crop TreeMean Crown Diameter on Cardinal DirectionsTree Quality (revised from Putnam 1960: preferred, reserve, or cutting)Number of Epicormic Branches per 16-foot logPoint System Hardwood Crown ClassesA point sample also was taken 2 feet to the west of each crop tree using a 10-factor prism. The following data were collected for all "in" trees:Azimuth to the treeDBHTotal HeightHeight to Green CrownDistance from the Crop TreeCrown Projection toward the Crop Tree All potential crop trees have been graded with a Crown Point System (Meadows, et al.), a score based on relative crown position, specifically: 1) the amount of crown receiving sunlight from directly above (scoring from 0, if completely overtopped, to 10, if completely free to grow), 2) how much of the top 50% of the crown received sunlight from the side (0-10), 3) crown balance (1-4), and 4) crown vigor (1-4). Crop trees must, 1) accumulate a Crown Point Score above 10, 2) have the potential to produce at least a 16-foot merchantable stem based on the existing stem and 3) not be a super dominant tree or be in an older age class. Obvious defects, such as sway, forks and excessive epicormic branching, will be noted and, if possible, avoided (Buckner and Houston 1997).Ice Storm One year after treatments a significant ice storm hit the mid-south. A number of crop trees were damaged, some totally destroyed, and quantitative individual-tree damage assessments were made soon after the storm. Three other studies have been initiated on the Ames Plantation, one in a 30-year-old bottomland stand concentrating on sweetgum (Liquidambar stryaciflua L.) and cherrybark red oak (Quercus falcata pagodaefolia Ell., a second in a 16-year-old bottomland clearcut focused on green ash (Fraxinus pennsylvanica Marsh.) and a third on an upland oak stand (Quercus sp.). Crop tree enhancement studies currently at Ames Plantation bring the total number of crop trees under treatment to about 1,750.Efforts Two Master's theses have reported the results of this study since treatments were applied. First, a Master's thesis was completed in 1995 by Dwight O'Neal. Significant gains in growth were reported. Twillman (2002) conducted a financial analysis and showed triple growth gains for release and fertilization with economic advantages held throughout the life of the stand. Another Master's project is currently underway. These studies will support future graduate student projects, one currently underway, and will lead to scholarly publications. They also will serve as the basis for extended economic evaluations as the trees continue to increase in size and value. The eventual body of knowledge will be available in University of Tennessee Extension publications. The research has been featured in forest landowner Field Days, workshops for professionals and as a teaching backdrop to illustrate silvicultural alternatives to forestry students.

Progress 03/16/15 to 01/31/19

Outputs
Target Audience:The target audience is 2-tiered. The first would be land managers, either the owners or professionals who makedecisions on howsilvicultural practices will be implemented. The second would be within academia, toscientists and also to those who dispense information to the public via extension of other means. The information is comprised with reasonable scientific evidence that planting of highly selected hardwood seedlings in post-harvest conditions can be accomplished successfully. And, at mid-rotational ages, the growth of these trees can be increased with release and fertilization treatments. Withpublication or on-site visits,information is being placed before these audiences and, as with any new scientific devlopment, being adopted as increasingly definitive results rise out of long term, complex and widely-arrayed studies. Changes/Problems:These are long-term studies and provideincreasingly strong results as they progress. What opportunities for training and professional development has the project provided? Nothing Reported How have the results been disseminated to communities of interest?Opportunities have included on-site field days for public and professional audiencess, incorporation of the research into formal collgiate coursework, and, with data collections underway, dissemination through scientific publication. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

Impacts
What was accomplished under these goals? It is perhaps safe to say among plant communities that are also widely utilized as commodities, hardwoods are essentially the last to be considered for domestication; and while this is not surprising, domesticationis also an inevitability. Hardwoods are comprised of a widely dispersed andcomplex set of species, some similarin terms of how and where they grow;but even with the subtleties of global marketing aside,they are very specific and very sensitive to species:site associations and bound to fail if these are not considered in their management. This wide array of specificrequirements, along with a near universal ability to sprout and retain genetic lineages across several generations, make them hard to work with.However, the domestication of pine, and especially loblolly(Pinus taeda)representedone of the moresuccessfulscientific efforts ever achieved and with it the theoretical groundwork was laid for similar work in hardwoods. And that work is needed, particularly in regeneration systems. The problems and uncertainties in the historicaldependence on natural regeneration in hardwoods, where adequate seed-in-place or an existing understory of desirable sapling-sized trees must be established and counted on, are hard to anticipate and success is difficult to predict. Once a stand full of undesirable trees is up and growing, alternatives to achieve a desirable mature stand can be limited by the species mix. It is a problem that canpersist throughout the life of the stand, often measured in decades.Some very desirable hardwoods, often the oaks (Quercus sp), are declining in regional and national inventories due to the vagaries of certain, increased diseases but also to an inability to successfully regeneratein the presence of an early-successional suite of species, a tree community that is very vigorous and typically predominated by sycamore, sweetgum and yellow-poplar (Platanus occidentalis, Liquidambar styraciflua, Liriodendron tulipfera, resp.). And, unfortunately because of the severity of naturally occuring competition, this is usually more-so true on the best sites.This projectis laying the foundation for domestication of hardwoods,especially the oaks, by successful establishment of seedlings in natural systems where a stand has been removed in a commercial operation and the resulting naturally occurring species mix is predictable only in terms of its competitive vigor, which is generally labelled as being "extremely high." However, its commercial and ecological value is less certain because it is overly occupied with less desirable species. Through a set of grading parameters we have come to understand the increased potential of certain root conformations and seedling size that will allow artificial regeneration to be competitive. An important concept has developed as we recognized the differences between pine plantations and mixed hardwood stands. Thatconcept is to "enrich" a developing stand, i.e., to establish enough predictably competitive seedlings to ensure the mature stand will have anenhanced component. In fact, on large ownerships or at landscape scales,"mixed stands" are desirable. Having an array of species contributes to ecological stability,wildlife values, andaesthetics. Along with that, a "portfolio" of merchantable opportunities gives the ability to respond to markets as they wax and wane, opportunities that might rangefrom cross ties to fine furniture, with sweetgum providing the first and oak the second. The research also is establishing the potential of a subset of the established trees to respond to mid-rotation treatments of release and fertilization, tripling growth in some cases. The core of the project'ssuccess is two-fold: first, artificially establishedoaks are showing free-to-grow tendencies at ages representing reasonable juvenile-to-mature predictions they willoccupythe mature stand, and second, there has been as much as a tripling ofgrowth in mid-rotational oaks where release and fertilization treatments were applied. These two developments, combined, indicate that oaks can be planted on very productive sites following a regeneration harvest, i.e., a clear-cut,a sub-set will enrich the ensuing stand, and the more successful can receive treatments that will increase productivity and decrease rotation length. And, this can be accomplished, and is most soundly accomplished, in the midst of a mixed stand. The study has documented the ability to establish "advance regeneration" by plantingthese pedigreed and highly selected seedling in single tree gaps where increased sunlight has reached the forest floor. This provides a way to establish regeneration in highly layered stands, usually occurring on very productivebottomland sites. A typical silvicultural treament in these stands, stands that are often bereft of hardwoodregeneration,is to kill the midstory to release light to the forest floor in hopes ofdeveloping an understory component with desirable seedlings. The project is showing the potential to follow a partial harvest with planting and releasing the artificially established trees later with a reproduction harvest. Within a mixed stand, every crop tree exists in the midst of a "competition cluster." That surrounding group will have considerable impact on a planted seedling's ability to survive and as it develops its status as a crop tree. For every seedling planted in this study,thecluster has been documented and in time, as the stand develops, there will be ancillary effects within acluster as treatmentsare applied to individual crop trees.Some of thesurrounding trees can be removed and the remaining trees responding as if in a thinning. Even where fertilization is applied to the crop tree, the neighbors can, to some extent, profit. Although these are long-term studies with increasing strength as they continue to develop, results are showing impressive survival rates, growth differences among species, and with these pedigreed seedlings, differences that are tending toward significance as the trees age. The concept of "enrichment plantings"is novel and presents a new way to evaluate plantings by giving weight to the best performers as opposed to averages derived across the entire study. For example, in a particular study where 200 trees per acre are planted, the average height across all trees at yeartenmight be tenfeet and the surrounding competition 25 feet. However, the best 25% of the planted trees might also average 25 feet, or better, and that subset of the original planting are now considered as free-to-grow trees (if the crown grading system, based on a 1-to-28 score, is high enough). These "winners" arecompetitively positioned to indicate probableoccupation in the mature stand. And, 25% of the original 200 would represent 50 trees/acres, well above the 36 trees/acres we have determined to represent a fully enriched stand, indeed, 36 mature oak would predominate an acre.The study is consistently showing results similar to the example, ranging from 20-to-100 free-to-grow trees at an age where juvenile: mature correlations are reasonably high. Even a 20-tree enrichment is not a "failure." For such a long-term investment as a full rotation, 20 high grade oak where none would exist otherwise is a good investment. In several settings where the project's findings have been presented, a general consensus among scientists and professionals, based on comments, has been a perception of this being "new science ... cutting edge ... ora turning over of old assumptions."

Publications


    Progress 10/01/17 to 09/30/18

    Outputs
    Target Audience: Nothing Reported Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided? Nothing Reported How have the results been disseminated to communities of interest?Forestry students in the Fall Block, Silviculture 322 course see the work in its entirety. The Fayette County Forestry Association centered its Annual Meeting around the research poject. What do you plan to do during the next reporting period to accomplish the goals?Alison Shimer, graduate student is scheduled to finish her project and manuscripts should be accomplished or be underway. John Bowers has manuscripts underway.

    Impacts
    What was accomplished under these goals? A Graduate Student, John Bowers, accomplished a Master's Thesis in December, 2016, "Crop Tree Enhancement of Green Ash (Fraxinus pennsylvantca) in a West Tennessee Bottomland;" and manuscritps are in process for publication in appropriate journals. Alison Shimer has been extended in her Master's Program to allow completion of her thesis reporting survival and growth of planted hardwoods. In that study eight species were established on small harvest openings, ranging from 1-to-3 acres. Ten years after establishment 75% (3,139) of the trees were alive. Several novel concepts have come from the work, incuding the defintion of successful regeneration defined as "enrichment" of a deloping stand with artificial regeneration but also maintining the economic and ecological values of a mixed stand. The basic goal is centered around having 36 planted trees per acre embeded in the mature, naturally occuring, mixed stand. Plantations are not expected or desired due to a suite of values associated with mixed stands and the unanticipated risks that can particularly focus on pure stands, with the emerald ash borer being a prime example. Along these lines, this project's developent suggests that research results presenting averages for all planted trees may be misleading when the top half or even just the top quarter of the planted population can be expected to occupy the mature canopy. In simple terms, prediciton of success is based on the "winners," i.e., the successful early-rotation competitors, defined with quantifiable criteria, and most likely and also in general desirably composed of a subset of all planted trees.

    Publications


      Progress 10/01/16 to 09/30/17

      Outputs
      Target Audience: Nothing Reported Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided? Nothing Reported How have the results been disseminated to communities of interest?A presentation was made at the Oak Symposium in Knoxville Tennessee, October 23-26, 2017. More than 150 land managers and scientists were in attendance and were able to hear the results of the project. A unique outcome from the meeting, beyond reporting the new concepts involved in this research, was the idea successfully planted stock can increase the agility needed for research designed to anticipate the biological and management adaptations required to deal with climate change. What do you plan to do during the next reporting period to accomplish the goals?Alison Shimer, graduate student is scheduled to finish her project and manuscripts should be accomplished or be underway.

      Impacts
      What was accomplished under these goals? A Graduate Student, John Bowers, accomplished a Master's Thesis in December, 2016. The work reported treatment response in a 30-year-old clearcut where green ash was primarily chosen for crop trees and received fertilization or release treatments. The research indicated significant growth gains and determined the point classification system developed by Meadows et. al. to be an important indicator for individual tree response potential. The research determined co-dominant trees are the most likely to respond to treatments. Additionally, soil analysis collect around each of 375 crop trees and primarily centered on mottling in the surface horizons indicated green ash grew best on the drier sites (mottling below 24 inches), but also these slightly drier sites supported a species mix that was somewhat less competitive and less adapted to these sites. Manuscripts are underway. Alison Shimer, a Master's Student, is near completion of a thesis reporting survival and growth of planted hardwoods, eight species, on small harvest openings that range from 1-to-3 acres. The study was established with 4,190 seedlings in 2006 and 2007. Ten years later 75% (3,139) were alive. Several primary results are apparent, including: sufficient free-to-grow crop trees are available to project mature correlations for a significantly enriched mature stand and as the plantings are maturing an increasing percentage of planted trees are reaching a free-to-grow status, exhibiting a negative-positive growth pattern where, for whatever reason, some planted trees stop growth and begin again as something happens inside their competition cluster to release them or the trees simply renew growth after building below-ground resources.

      Publications


        Progress 10/01/15 to 09/30/16

        Outputs
        Target Audience:Scientists, professional land managers, and landowners. Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided?Undergraduate Forestry Students from the University of Tennessee spend a day in the field seeing the entire range of the research. How have the results been disseminated to communities of interest?In the spring of 2016, 50 scientists and professional land managers from the Southern Region attended a two-day workshop centered on the Precision Forest Research. Senior scientists in that meeting commented this as being the most significant hardwood research nation-wide. What do you plan to do during the next reporting period to accomplish the goals?The research will continue as a tool for student education.

        Impacts
        What was accomplished under these goals? Poster: John Luke Bowers, Wayne Clatterbuck, Allan Houston, Donald Tyler, John Zobel, Evaluating Crop Tree Enhancement of Green Ash in a west Tennessee Hardwood Bottom from the Perspective of Crown Class, Southern Hardwood Forestry Group Annual Meeting, February, 2016, USFS, Stoneville, MS.

        Publications


          Progress 03/16/15 to 09/30/15

          Outputs
          Target Audience: The target audience for research output has been natural resource managers, timber consultants, forest conservationists, private landowners, biologists and those who design forest policy. Tour and research overview: regional landowners and professional foresters on-site with 55 people attending. Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided?A University of Tennessee, extension program was conducted on-site with 55 professional foresters and landowners in attendance. How have the results been disseminated to communities of interest?Tour and research overview: regional landowners and professional foresters on-site with 55 people attending. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

          Impacts
          What was accomplished under these goals? The research is centered on developing high quality hardwood stands with the enrichment of natural regeneration in small harvested openings with the addition of high quality seedlings and the improvement of growth during intermediate stages using fertilization and release. With attention to harvest, regeneration and intermediate stages, the work will develop a silvicultural set of treatments that can be alpplied over the lifetime of a hardwood forest. A large number of group selection openings of 1-to-3 acres have been established over the past 10 years in various portions of the Ames Plantation Forest. These openings have been planted with highly selected hardwood seedlings, primarily several oak species, but also including walnut. Seedlings are planted on a wide spacing, often 20'X20', with a goal of having a subset persist into the upper canopy. If as few as 25% of the planted stock are in a free-to-grow situation at the end of 6 years it can indicate eventual success because 25 large oak at maturity can dominate an acre. On many of these sites and representative of many harvested openings, including clearcuts, oak is often missing in the natural regeneration unless it was established as advance regeneration. A Masters project was begun in 2015 to evaluate survival and growth of the planted trees in harvested openings on 9 bottomland sites. Across all sites, 4,191 seedlings composed if 7 oak species and black walnut from 88 genetic families were established in an incomplete block design. Six sites were planted in 2006 and 3 in 2007. Measurements conducted in 2013 found tree averages on some sites ranging upwards of 5 meters. These 9 sites will form the basis for the Master's project. In all there are more than 20,000 trees in the precision research. Another Masters project was initiated on a Crop Tree Enhancement project composed of 375 green ash trees resulting from a clearcut conducted in 1980. At 14-years-of-age the resulting stand was assessed for viable crop trees and due to an unusual component of green ash, the project was focused on the response of this species to release, fertilization and a combination of the two treatments. The Masters project will examine growth response, and additionally will provide insight into which segment of the crop tree population is most viable for treatment response. Field measurements were completed in the summer of 2015, including a soil sample taken around every tree. Preliminary analyses suggest the lower third, based on initial measurements, of crop trees were unable to respond adequately to treatments. The upper third did not appreciably respond. However, the middle third responded to treatments and improved their relative stature in the stand, indicating there is a target condition within these stands where treatments can be most effectively applied. The crop tree work extends over several sites and several species. As means come available, we will examine these species for similar trends. Some crop tree studies are being evaluated for a repetition of treatments. A Masters study completed in 2002 indicated as much as a tripling of growth with release and fertilization treatments and the significant response holding for 40 years beyond application.

          Publications

          • Type: Conference Papers and Presentations Status: Other Year Published: 2015 Citation: Bowers, J.L., Clatterbuck, W., Houston, A.E., Zobel, J., Tyler, D. (2015, November). The Impact of Pre-Treatment Diameter on the Growth of Green Ash Crop Trees in a West TN Hardwood Bottom. Poster presented at the 2015 Society of American Foresters National Convention, Baton Rouge, LA.