Source: NORTH CAROLINA STATE UNIV submitted to NRP
ENHANCING THE CUCURBIT DOWNY MILDEW FORECASTING SYSTEM
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
0221747
Grant No.
2010-41530-21134
Cumulative Award Amt.
(N/A)
Proposal No.
2010-02685
Multistate No.
(N/A)
Project Start Date
Jul 1, 2010
Project End Date
Jun 30, 2013
Grant Year
2010
Program Code
[QQ.S]- Integrated Pest Management - South Region
Recipient Organization
NORTH CAROLINA STATE UNIV
(N/A)
RALEIGH,NC 27695
Performing Department
Plant Pathology
Non Technical Summary
The U.S. grows about 466,250 acres of cucurbits valued at $1.7 billion. In 2004, severe outbreak of the disease occurred in the eastern U.S., devastating cucurbit crops. Many fields were abandoned without harvesting and suffered 100% loss. Fungicide programs initially failed either because applications were made after disease was established, and/or ineffective products were used. Resistance of Pseudoperonospora cubensis to specific fungicides was suspected as a contributor to the ineffectiveness of fungicide programs. A forecasting system was developed to help growers to make decisions on if and when to spray based on disease outbreaks in sentinel plots located in 25 states and Ontario, Canada. The ultimate goal is to automate the current forecasting system and minimize the input of human element. However, a series of research projects to develop pathogen biology models is needed before the forecasting system can be fully automated. We propose to conduct the research necessary to enable growers to improve cucurbit downy mildew control, reduce occurrence of ineffective fungicide applications, and implement improved resistance management programs. This project will focus on enhancement of the disease forecasting system, documentation of the presence/absence of differential fungicide resistance in the eastern U.S. and establish the population structure of P. cubensis within the sentinel network. We will continue to work with stakeholders to enable growers to reduce costly and unnecessary fungicide sprays, reduce risk of resistance development to fungicides and improve chemical control of cucurbit downy mildew.
Animal Health Component
60%
Research Effort Categories
Basic
(N/A)
Applied
60%
Developmental
40%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
21614291170100%
Knowledge Area
216 - Integrated Pest Management Systems;

Subject Of Investigation
1429 - Cucurbits, other;

Field Of Science
1170 - Epidemiology;
Goals / Objectives
Beginning January 2010, current funding for the Cucurbit forecasting system will expire for the sentinel monitoring network for reporting disease outbreaks via the website and as of October 2010, current funding will expire for the entire project. Currently, collaborators in the sentinel network depend on the cucurbit downy mildew forecasting system to serve their clientele efficiently. Given the importance of the already established network in delivering service to cucumber growers in the eastern U.S., California and Ontario, Canada, the 25 collaborating institutions on the Cucurbit ipmPIPE have agreed to continue with sentinel monitoring and reporting of disease outbreaks as part of their regular extension activities, at their own expense at an estimated cost of $100,000 per year. Likewise, our programs and those of our collaborators will continue to screen fungicides and other control practices to improve control recommendations. However, the backbone of effective development of recommendations, dissemination of information and timely disease forecasts relies on an effective website and program coordination. There is no other institution or organization that currently provides growers with downy mildew forecasts on the scale of the Cucurbit ipmPIPE project. Integrated management of cucurbit downy mildew is based on the use of sound IPM principles, including host plant resistance, disease forecasting and the preventive use of effective fungicides. Project cooperators are investigating these inter-connected disease management strategies, and have chosen to focus this proposal on filling critical and timely research needs. Specifically, we propose to conduct the research necessary to enable growers to improve cucurbit downy mildew control, reduce ineffective fungicide applications, and implement improved resistance management programs. Our objectives will be: i) refine the cucurbit downy mildew disease forecasting system, and ii) establish the population structure of P. cubensis population in the sentinel network in eastern United States.
Project Methods
The enhancement of the downy mildew forecasting system will be accomplished through two sub-objectives: i) Quantitative relationship between disease severity and source strength. Experiments will be conducted in two locations, Clayton and Clinton, North Carolina during the summer. The susceptible cucumber variety will be used at both locations in 22 rows, 100 m long plots. Sites will be selected carefully to ensure that the area surrounding plots will be fairly flat and no obstacles to wind movement are present in the surrounding area. Cucumber leaves from plants inoculated will be used to inoculate plots 4 weeks after planting. Inoculation will be accomplished by scattering the infected leaves in a 5-m diameter circular area of cucumber plants. To protect plants within the source area from infection outside the source area, Presidio will be applied to the un-inoculated area before inoculation. Disease severity will be monitored periodically and the following variables will be determined: a) standing crop of sporangia in the source, and b) airborne sporangia concentration. Each morning, before the dew dries from the plant surfaces, four sites within a 5-m diameter circle will be sampled to obtain the number of sporangia per unit of ground area within the canopy. The total number of lesions within a 0.25-m2 grid will be counted at each of the four sites in the canopy. At each of the four sites, three to five lesions will be destructively sampled to assess sporulation per unit area. Aerial concentrations of P. cubensis sporangia will be measured above the spore sources using Rotorod spore samplers with retracting-type sampling heads. The Rotorods will be operated for 15-20 min every hour for each sampling period. At the end of each sampling period, the sampling rods will be replaced and a new sampling period begun. Eight 1-h periods (8 h of sampling) will be carried out on each day. Counts of sporangia on the Rotorods will be converted to spore concentrations by accounting for the proportion of the sample surface that was counted, the sampling rate, and the duration of the sampling period. ii) Effectiveness of FLEXPART model. Currently, spore transport in the atmosphere is calculated using the HYSPLIT trajectory model from the National Oceanic and Atmospheric Association's (NOAA) Air Resources Laboratory (ARL). The data of primary interest are the forecast wind fields in the atmospheric boundary layer. HYSPLIT trajectories are provided by ARL via a restricted-access website. The trajectory is a plot of the future atmospheric pathway of a "parcel" of air likely to contain spores; in other words, the prediction of the spatial and temporal positions of a spore cloud center for the next two days following release from a source site. Here, outputs from the HYSPLIT will be compared to the FLEXPART model to determine the similarity in the predictions. The FLEXPART model is particularly attractive since it has a finer resolution and effects of environmental conditions and source strength can easily be incorporated in the model.

Progress 07/01/10 to 06/30/13

Outputs
Target Audience: 1. Growers that are producers of cucumbers, melon, squah, pumpking and watermelon located in across the country where cucurbits are grown. Growers signed up to recieve diseasealerts for the risk of cucurbit downy mildew within their area of operation. 2. The project also focused on crop extension specialists, extension agents and crop consultants that regularly have cucurbit operations during the growing season. Changes/Problems: Nothing Reported What opportunities for training and professional development has the project provided? 1. Traning: Katie Neufeld received herMS degree from the Department of Plant Pathology, North carolina State University in 2012. How have the results been disseminated to communities of interest? Results were disseminated to the growers at stakeholders by state colloborators on the project through several means: 1. Brochures. 2. Forecasting website (http://cdm.ipmpipe.org) 3. Growers meeting and professional societal meeting such as American Phytopathological Society Meeting and Pickle Packers International. 4. Farmer field days and extension meetings. What do you plan to do during the next reporting period to accomplish the goals? Nothing Reported

Impacts
What was accomplished under these goals? The cucurbit downy mildew forecasting system was refined in a number of ways: 1. The Flexpart model was adopted as the plume transport model topredict the trajactory pathway for spore transport from source fields. 2. The forecasting system was made to be biologicallysound by incorporating the relationship between disease severity at the source field and the amount of spores of the pathogen that can be released. 3. Models of combined effects of temperature and leaf wetness duration on disease severity were used to delineate areas of different levels of disease risk based on forecasted temperature and leaf wetness duration.

Publications

  • Type: Journal Articles Status: Published Year Published: 2013 Citation: Neufeld, K.N., Isard, S.A., Ojiambo, P.S. 2013. Relationship between disease severity and escape of Pseudoperonospora cubensis sporangia from a cucumber canopy during downy mildew epidemics. Plant Pathology 2013. Doi: 10.1111/ppa.12040.
  • Type: Journal Articles Status: Published Year Published: 2012 Citation: Neufeld, K.N., and Ojiambo, P.S. 2012. Interactive effects of temperature and leaf wetness duration on sporangia germination and infection of cucurbit hosts by Pseudoperonospora cubensis. Plant Disease 96:345-353.
  • Type: Theses/Dissertations Status: Published Year Published: 2012 Citation: Neufeld, K.N. 2012. Epidemiology of Cucurbit Downy Mildew: Effects of Weather Variables on Infection Parameters and Disease Severity and Source Strength Relationships.


Progress 07/01/10 to 06/30/11

Outputs
OUTPUTS: Field experiments were conducted at two sites in North Carolina namely; Clayton (Johnston County) and Clinton (Sampson County) to determine the relationship between the severity of cucurbit downy mildew and the amount of P. cubensis sporangia produced and proportion of sporangia that escaped the canopy. The experiments were performed using the cucumber cultivar Poinsette 76 and plants relied on natural infection. Disease severity was monitored during the summer and assessed visually based leaf area infected. At disease levels of 1, 5, 15, 20, 30 and 40%, total standing sporangia was estimated based on lesions with a 0.25 by 0.25 m grid and sporulation per unit area. Aerial concentration of sporangia was quantified using Rotorod spore samplers mounted at 0.5, 1.0 and 1.5 m above the canopy. Concentrations were monitored from 7 to 3 pm on each level of disease severity. Counts of sporangia on the Rotorods were then converted to spore concentrations by accounting for the proportion of the sample surface that was counted, the sampling rate, and the duration of the sampling period. The relationship between standing crop of sporangia in the source and airborne sporangia concentration at different heights above the canopy and disease severity was analyzed using PROC REG in SAS. Disease forecasting data were gathered from previous forecasting output when HYSPLIT model was used to predict spore transport. A similar set of data is being gathered based on FLEXPART as the model for spore dispersal. PARTICIPANTS: 1. Katie Neufeld, Graduate Student working on the epidemiology and management of cucurbit downy mildew. 2. Mike Adams, lab manager who worked and coordinated activities of the graduate student in the lab including field layout and data collection. 3. Rob Kautz, undergraduate summer student that assisted with field experiments, 4. Wendy Britton, project coordinator, and 5. Thomas Keever, forecaster who both worked to compile data to compare the effectiveness of the HYSPLIT versus the FLEXPART models in predicting trajectories of sporangia transport. TARGET AUDIENCES: 1. Cucurbit growers, 2. Extension county agents, 3. Industry specialists PROJECT MODIFICATIONS: Nothing significant to report during this reporting period.

Impacts
Results of sporangia concentrations indicate that across all levels of disease severity, peak sporangia production occurred between 8 and 10 am and then decreased thereafter. The concentration of sporangia also decreased linearly with increasing height above the canopy with the concentration being highest at 0.5 m above the canopy and lowest at 1.5 m above the canopy. Sporangia concentration at 1.0 m above the canopy was intermediate. These results were similar for both experiments conducted at Clayton and Clinton. Graphical results of the relationship between disease severity and sporangia concentration indicate that the relationship is best described by a gamma function with concentration increasing with increasing severity up to 20% and then tailing off as disease severity increases.

Publications

  • No publications reported this period