Source: AUBURN UNIVERSITY submitted to NRP
ASSESSING VARIATIONS IN FEED EFFICIENCY OF ANGUS-SIMMENTAL CATTLE: APPLICATION OF FUNCTIONAL GENOMICS/GENE EXPRESSION ANALYSIS IN A LONG TERM BREEDING STUDY
Sponsoring Institution
National Institute of Food and Agriculture
Project Status
COMPLETE
Funding Source
Reporting Frequency
Annual
Accession No.
0209038
Grant No.
(N/A)
Cumulative Award Amt.
(N/A)
Proposal No.
(N/A)
Multistate No.
(N/A)
Project Start Date
Oct 1, 2006
Project End Date
Sep 30, 2007
Grant Year
(N/A)
Program Code
[(N/A)]- (N/A)
Recipient Organization
AUBURN UNIVERSITY
108 M. WHITE SMITH HALL
AUBURN,AL 36849
Performing Department
ANIMAL SCIENCES
Non Technical Summary
Efficiency of performance is paramount for the continued profitability and sustainability of the US Beef Cattle Industry. In the past, production efficiency has been evaluated by relating performance to output, typically total or rate of weight gain. However the fastest gaining animals are not necessarily the most efficient since they may exhibit high maintenance costs. It has been shown that feed efficiency (FE) and a related measure, residual feed efficiency (RFI) are more highly related to efficiency of performance and would be useful as more robust selection criteria in breeding programs. Unfortunately FE and RFI can only be determined by a long term feeding trial for each individual animal. Thus determinations of FE and RFI is cost prohibitive and hence the use of these efficiency parameters in breeding programs by the commercial beef cattle industry is severely limited. The purpose of this project is to develop a process, based on expression of genes related to performance efficiency in fat and muscle tissues of cattle, that can reliably predict which animals will have better FE and RFI. It has already been shown that FE and RFI in cattle are moderately heritable and amenable to selection and improvement via well designed breeding programs. The use of expression patterns of specific efficiency marker genes measured while animals are still young, can then be utilized as more robust selection tools for superior bulls and females and subsequent mating strategies.
Animal Health Component
10%
Research Effort Categories
Basic
80%
Applied
10%
Developmental
10%
Classification

Knowledge Area (KA)Subject of Investigation (SOI)Field of Science (FOS)Percent
3023310104030%
3023310108010%
3033310104040%
3033310108010%
3043310104010%
Goals / Objectives
The objective of this one year study is to test the hypothesis that in cross bred beef cattle expression of patterns of genes involved in oxidative metabolism, lipid anabolism and protein turnover will be related to variations in feed efficiency among similar sized animals in the late growing phase.
Project Methods
At the Alabama Agricultural Experiment Station Black-Belt Substation a cattle breeding /selection program was initiated to develop strategies to select cattle for residual feed efficiency (RFI). Progeny of matings of sires and dams selected for higher and lower potential RFI (16 total cross bred cattle) after weaning at 325 Kg will be assigned to a 84 day feeding trial; daily feed intake and weight gain will be measured. On day 42, tissue biopsies will be secured (September, 2006) from tail head adipose depots and from loin muscle. All tissues will be powdered cryogenically and RNA will be isolated using the Trizol procedure. Harvested RNA will then be purified and gene expression will be accomplished with quantitative RT-PCR using a 2 step (RT first) approach. M-RNA abundance will be determined for the following genes in 1) skeletal muscle: PPAR α, cyto-C oxidase, CPT-1, ubiquitin conjugating enzymes E2, ubiquitin specific protease, UCP2 and 3; in 2) adipose tissue: PPARγ2, fatty acid synthase, leptin, GPAT, UCP 2 and 3 and PCG-1α. From the feeding trial, the RFI for each animal will be determined; the gene expression results for each animal will then be related to RFI with appropriate regression and statistical procedures to test for the significance of correlation between RFI and fatty acid oxidation, synthesis, mitochondrial efficiency and protein turnover.

Progress 01/01/07 to 12/31/07

Outputs
This is a one year project but was not funded until 5 month into the 12 month project period. Thus we have just completed the animal work and are now measuring gene expression of various genes related to efficiency of cattle production. There are NO data yet but the work will be completed this calendar year.

Impacts
No outcome anfd impacts yet.

Publications

  • No publications reported this period


Progress 10/01/06 to 09/30/07

Outputs
OUTPUTS: As noted last year the funding for this approved project came too late to write an appropriate report for 2007. The aim of this project was to relate expression of selected genes in skeletal muscle and adipose tissue of finishing steers to performance efficiency measures as feed efficiency and residual feed index. The results for the completed study are as follows: Metabolic redundancies and inefficiencies at the molecular level associated with feed efficiency (FE) and residual feed intake (RFI) of food animals may be ascertained by gene expression (GE) profiling. The objective of this study was to examine relationships between GE (mRNA abundance) of fatty acid synthase (FAS), PPAR α, PPAR γ2, carnitine palmitoyl transferase (CPT-1b), leptin, uncoupling protein 2 (UCP-2), ubiquitin conjugating enzyme (E2) and polyubiquitin (PQ) genes in skeletal muscle (SKM) and adipose tissue (AT) with FE and RFI in finishing cattle. Crossbred cattle (bulls, 8; steers, 8) were placed on a feedlot diet and AT and SKM biopsies were taken prior to completion of the finishing trial. Gene expression was determined with real time RT-PCR. Adipose FAS, leptin and PPARγ2 GE were unrelated to RFI. SKM UCP-2 and E-2 but not CPT-1b, PPARα and PQ GE were correlated with RFI (P<.05). AT leptin GE was higher (P<.05) while SKM UCP-2 GE was lower (P<.05) in bulls than steers. While PQ GE was not correlated with RFI it was highly correlated (P<.01) with E2 GE. Results indicate that AT genes for fat deposition are likely minimally related to RFI and FE, while SKM genes associated with mitochondrial metabolism and protein turnover are correlated with RFI. The transcriptomic signature (GE) for efficiency in beef cattle appears more related to SKM than AT metabolism. PARTICIPANTS: Not relevant to this project. TARGET AUDIENCES: The results of this work provide the following guidance for the future. 1. A transcriptomic/gene expression signature for individual beef cattle determined by quantitative RT-PCR of skeletal muscle, adipose and liver biopsis will be an effective procedure to conduct a performance/phenotypic evaluation of individual cattle particularly in grazing systems. 2. This type of work not only will increase our understanding of mechanisms at the genomic/molecular regulation level that impinge on production efficiency, but such results can be used to design improved beef cattle feeding strategies for grazing systems in the Southeast. Finally these type of data may be useful in deciding whether performance selection based on sire evaluation of progeny in the feedlot will be appropriate for selection of cattle most productive in grazing systems. PROJECT MODIFICATIONS: Not relevant to this project.

Impacts
This project was a beta test to investigate whether gene expression analysis from beef cattle biopsies (obtained during various time points of the production phase) may be utilized to assess performance efficiency independently of measuring traditional parameters such as feed intakes, daily gain and calculations of feed/gain ratios (a measure of performance efficiency including maintenance needs of animals). Our data revealed that there is a gene expression signature that relates to beef cattle performance. This signature indicated that adipose metabolism is neutral to performance efficiency while mitochondrial function and protein degradation/remodelling in skeletal muscle will have significant effects on performance. These results would be supportive of the idea that poorly performing cattle exhibit greater skeletal muscle protein remodeling and uncoupling of ATP production. This is the first of such studies and additional work is needed to move these ideas from the hypothesis stage to an application stage in animal selection and feeding/management regimes.

Publications

  • No publications reported this period